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Plant pathology
Plant pathology












plant pathology

Preface xxi Photo credits xxiii About the Author xxvi Finally, to Mark and Maximos, our youngest grandchildren, who, someday, when they read their names in the book, may be reassured of “Granpa’s” love for them, and may feel proud of their grandfather. To my daughters-in-law, Betsy and Vivynne, who, by joining our family, added beauty, love, enjoyment, and four wonderful grandchildren. To my wife, Annette, whose love and support have been the most precious things to me throughout our life together, and who helped me in many facets of preparation of this and of previous editions of Plant Pathology.

plant pathology

To my sisters, Dimitra and Evangelia, who have been there for me forever and who also sacrificed some of their interests for my benefit. Buchholtz, my major professor at Iowa State University, who challenged me before I had even taught my first lecture to “write my own textbook on Plant Pathology”. This, the 5th and probably the last edition of Plant Pathology by me, is dedicated: To the memory of my parents, Nikolas and Olga, who, in spite of their limited education, sacrificed everything to give me the most and best education possible. SB731.A35 2004 571.9¢2 - dc22 2004011924 British Library Cataloguing in Publication Data A catalogue record for this book is available from the British Library For all information on all Elsevier Academic Press Publications visit our Web site at Printed in the United States of America 04 05 06 07 08 09 9 8 7 6 5 4 3 2 1 Includes bibliographical references and index. Permissions may be sought directly from Elsevier’s Science & Technology Rights Department in Oxford, UK: phone: (+44) 1865 843830, fax: (+44) 1865 853333, e-mail: You may also complete your request on-line via the Elsevier homepage (), by selecting “Customer Support” and then “Obtaining Permissions.” Library of Congress Cataloging-in-Publication Data Agrios, George N., 1936– Plant pathology / George Agrios. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system, without permission in writing from the publisher. Sonnack Troy Lilly Linda Beattie Eric DeCicco SNP Best-set Typesetter Ltd., Hong Kong RR Donnelley & Sons Company RR Donnelley & Sons CompanyĮlsevier Academic Press 30 Corporate Drive, Suite 400, Burlington, MA 01803, USA 525 B Street, Suite 1900, San Diego, California 92101-4495, USA 84 Theobald’s Road, London WC1X 8RR, UK This book is printed on acid-free paper. Publisher Associate Acquisitions Editor Project Manager Marketing Manager Cover Design Composition Cover Printer Interior Printerĭana Dreibelbis Kelly D. AGRIOS Department of Plant Pathology University of Florida














Plant pathology